Coyote
Drysophila
“Two strains of Drosophila paulistorum developed hybrid sterility of male offspring between 1958 and 1963. Artificial selection induced strong intra-strain mating preferences.
(Test for speciation: sterile offspring and lack of interbreeding affinity.) “
http://www.talkorigins.org/faqs/speciation.html
If I am reading this correctly, only the males are infertile to the parent species. Are the females still fertile to the parent species? This sounds like an infertility problem and not true speciation to me.
Not really - from what I understand, it's a form of mechanical isolation that forms one of the mechanisms for speciation to occur.
For example: this is from an abstract describing it describing speciation in another varient of drosophila (and I won't pretend I understand all the science here):
http://www.pubmedcentral.nih.gov/articlerender.fcgi?artid=42249
It is generally believed that Drosophila melanogaster has no closely related species with which it can produce the viable and fertile hybrids that are essential for the genetic analysis of speciation. Following the recent report of molecular differentiation between a Zimbabwe, Africa, population and two United States populations, we provide evidence that strong sexual isolation exists between the D. melanogaster population in Zimbabwe and populations of other continents. In the presence of males of their own kind, females from most isofemale lines of Zimbabwe would not mate with males from elsewhere; the reciprocal mating is also significantly reduced, but to a lesser degree. The genes for sexual behaviors are apparently polymorphic in Zimbabwe and postmating reproductive isolation between this and other populations has not yet evolved. Whole chromosome substitutions indicate significant genetic contributions to male mating success by both major autosomes, whereas the X chromosome effect is too weak to measure. In addition, the relative mating success between hybrid and pure line males supports the interpretation of strong female choice. These observations suggest that we are seeing the early stages of speciation in this group and that it is driven by sexual selection. The genetic and molecular tractability of D. melanogaster offers great promise for the detailed analysis of this apparent case of incipient speciation.
Faeroe Island Mouse
“The Faeroe mouse has been successfully crossed with the English house mouse“.
http://links.jstor.org/sici?sici=0021-8790(193811)7:2<290:NOTBOT>2.0.CO;2-H
The Faeroe Island Mouse can’t be a new species if it can mate with an existing species.
True.
Lake Nagubago ciclids“(Test for speciation in this case is by morphology and lack of natural interbreeding. These fish have complex mating rituals and different coloration. While it might be possible that different species are inter-fertile, they cannot be convinced to mate.)”
http://www.talkorigins.org/faqs/speciation.html
Based on the above, it appears to me the jury is still out on the Lake Nagubago ciclids. No speciation has been observed here, to date.
Incorrect. Mating behavior in fish and insects is genetically determined and complex - if they can not be convinced to mate, they are likely if not all ready there then well on their way to being different species.
Still no verified, undisputed speciation here in any of the examples you cited. Would you like to try again?
Sure - there quite a few listed here: mostly plants and insects.
http://www.talkorigins.org/faqs/faq-speciation.html
In addition, from the same source:
# New species have arisen in historical times. For example:
* A new species of mosquito, the molestus form isolated in London's Underground, has speciated from Culex pipiens (Byrne and Nichols 1999; Nuttall 1998).
* Helacyton gartleri is the HeLa cell culture, which evolved from a human cervical carcinoma in 1951. The culture grows indefinitely and has become widespread (Van Valen and Maiorana 1991).
A similar event appears to have happened with dogs relatively recently. Sticker's sarcoma, or canine transmissible venereal tumor, is caused by an organism genetically independent from its hosts but derived from a wolf or dog tumor (Zimmer 2006; Murgia et al. 2006).
* Several new species of plants have arisen via polyploidy (when the chromosome count multiplies by two or more) (de Wet 1971). One example is Primula kewensis (Newton and Pellew 1929).
# Incipient speciation, where two subspecies interbreed rarely or with only little success, is common. Here are just a few examples:
* Rhagoletis pomonella, the apple maggot fly, is undergoing sympatric speciation. Its native host in North America is Hawthorn (Crataegus spp.), but in the mid-1800s, a new population formed on introduced domestic apples (Malus pumila). The two races are kept partially isolated by natural selection (Filchak et al. 2000).
* The mosquito Anopheles gambiae shows incipient speciation between its populations in northwestern and southeastern Africa (Fanello et al. 2003; Lehmann et al. 2003).
* Silverside fish show incipient speciation between marine and estuarine populations (Beheregaray and Sunnucks 2001).
Anyway - we really would not expect to observe large changes directly. Evolution consists mainly of the accumulation of small changes over large periods of time - very large. We simply havent' been around (and aware) for long enough.